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On the Origin of Species

Charles Darwin's On the Origin of Species, published on 24 November 1859, is a work of scientific literature which is considered to be the foundation of evolutionary biology. Its full title was On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life. For the sixth edition of 1872, the short title was changed to The Origin of Species. HP Omnibook XE3 Battery

Darwin's book introduced the scientific theory that populations evolve over the course of generations through a process of natural selection. It presented a body of evidence that the diversity of life arose by common descent through a branching pattern of evolution. HP Omnibook XT1000 Battery

Darwin included evidence that he had gathered on the Beagle expedition in the 1830s and his subsequent findings from research, correspondence, and experimentation. HP Omnibook XT1500 Battery

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Various evolutionary ideas had already been proposed to explain new findings in biology. There was growing support for such ideas among dissident anatomists and the general public, but during the first half of the 19th century the English scientific establishment was closely tied to the Church of England, while science was part of natural theology. HP Pavilion DV2000 Battery

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Ideas about the transmutation of species were controversial as they conflicted with the beliefs that species were unchanging parts of a designed hierarchy and that humans were unique, unrelated to other animals. The political and theological implications were intensely debated, but transmutation was not accepted by the scientific mainstream. HP Pavilion DV2200 Battery

The book was written for non-specialist readers and attracted widespread interest upon its publication. As Darwin was an eminent scientist, his findings were taken seriously and the evidence he presented generated scientific, philosophical, and religious discussion. HP Pavilion DV2300 Battery

The debate over the book contributed to the campaign by T.H. Huxley and his fellow members of the X Club to secularise science by promoting scientific naturalism. Within two decades there was widespread scientific agreement that evolution, HP Pavilion DV2400 Battery

with a branching pattern of common descent, had occurred, but scientists were slow to give natural selection the significance that Darwin thought appropriate. During the "eclipse of Darwinism" from the 1880s to the 1930s, various other mechanisms of evolution were given more credit. HP Pavilion DV2500 Battery

With the development of the modern evolutionary synthesis in the 1930s and 1940s, Darwin's concept of evolutionary adaptation through natural selection became central to modern evolutionary theory, now the unifying concept of the life sciences. HP Pavilion DV2600 Battery

DEVELOPMENTS BEFORE DARWIN'S THEORY

In later editions of the book, Darwin traced evolutionary ideas as far back as Aristotle;[4] the text he cites is a summary by Aristotle of the ideas of the earlier Greek philosopher Empedocles.[5] HP Pavilion DV2700 Battery

Early Christian Church Fathers and Medieval European scholars interpreted the Genesis creation myth allegorically rather than as a literal historical account;[6]organisms were described by their mythological and heraldic significance as well as by their physical form. Nature was widely believed to be unstable and capricious, with monstrous births from union between species, and spontaneous generation of life.[7]HP Pavilion DV4 Battery

 

The Protestant Reformation inspired a literal interpretation of the Bible, with concepts of creation that conflicted with the findings of an emerging science seeking explanations congruent with the mechanical philosophy of René Descartes and the empiricism of the Baconian method. HP Pavilion DV4T Battery

After the turmoil of the English Civil War, the Royal Society wanted to show that science did not threaten religious and political stability. John Ray developed an influentialnatural theology of rational order; in his taxonomy, HP Pavilion DV5 Battery

 species were static and fixed, their adaptation and complexity designed by God, and varieties showed minor differences caused by local conditions. In God's benevolent design, carnivores caused mercifully swift death, but the suffering caused by parasitism was a puzzling problem. HP Pavilion DV5T Battery

The biological classificationintroduced by Carolus Linnaeus in 1735 also viewed species as fixed according to the divine plan. In 1766, Georges Buffonsuggested that some similar species, such as horses and asses, or lions, tigers, and leopards, HP Pavilion DV7T Battery

might be varieties descended from a common ancestor. The Ussher chronology of the 1650s had calculated creation at 4004 BC, but by the 1780s geologists assumed a much older world. Wernerians thought strata were deposits from shrinking seas, but James Hutton proposed a self-maintaining infinite cycle, anticipating uniformitarianism.[8]HP Pavilion DV4000 Battery

 

Charles Darwin's grandfather Erasmus Darwin outlined a hypothesis of transmutation of species in the 1790s, and Jean-Baptiste Lamarck published a more developed theory in 1809. Both envisaged that spontaneous generation produced simple forms of life that progressively developed greater complexity, HP Pavilion DV5000 Battery

adapting to the environment by inheriting changes in adults caused by use or disuse. This process was later called Lamarckism. Lamarck thought there was an inherent progressive tendency driving organisms continuously towards greater complexity, in parallel but separate lineages with no extinction.[9] HP Pavilion DV6100 Battery

Geoffroycontended that embryonic development recapitulated transformations of organisms in past eras when the environment acted on embryos, and that animal structures were determined by a constant plan as demonstrated by homologies. HP Pavilion DV6300 Battery

Georges Cuvier strongly disputed such ideas, holding that unrelated, fixed species showed similarities that reflected a design for functional needs.[10] His palæontological work in the 1790s had established the reality of extinction, which he explained by localcatastrophes, followed by repopulation of the affected areas by other species.[11]HP Pavilion DV6500t Battery

In Britain, William Paley's Natural Theology saw adaptation as evidence of beneficial "design" by the Creator acting through natural laws. All naturalists in English universities were Church of England clergymen, and science became a search for these laws.[12] HP Pavilion DV8000 Battery

Geologists adapted catastrophism to show repeated worldwide annihilation and creation of new fixed species adapted to a changed environment, initially identifying the most recent catastrophe as the biblical flood.[13] Some anatomists such as Robert Grant were influenced by Lamarck and Geoffroy, but most naturalists regarded their ideas of transmutation as a threat to divinely appointed social order.[14]HP Pavilion DV8002ea Battery

INCEPTION OF DARWIN'S THEORY

Darwin went to Edinburgh University in 1825 to study medicine. In his second year he neglected his medical studies for natural history and spent four months assisting Robert Grant's research into marine invertebrates. Grant revealed his enthusiasm for the transmutation of species, HP Pavilion DV9000 Battery

but Darwin rejected it.[15] AtCambridge University starting in 1827, Darwin learnt science as natural theology from botanist John Stevens Henslow, and read Paley, John Herschel and Alexander von Humboldt. Filled with zeal for science, he studied catastrophist geology with Adam Sedgwick.[16][17]HP Pavilion DV9100 Battery

In December 1831, he joined the Beagle expedition as a geologist and naturalist. He read Charles Lyell's Principles of Geology and from the first stop ashore, at St. Jago, found Lyell's uniformitarianism a key to the geological history of landscapes. HP Pavilion DV9500 Battery

Darwin discovered fossils resembling huge armadillos, and noted the geographical distribution of modern species in hope of finding their "centre of creation".[18] The three Fuegian missionaries the expedition returned to Tierra del Fuegowere friendly and civilised, HP Pavilion DV9600 Battery

yet to Darwin their relatives on the island seemed "miserable, degraded savages",[19] and he no longer saw an unbridgeable gap between humans and animals.[20] As the Beagle neared England in 1836, he noted that species might not be fixed.[21]HP Pavilion N3200 Battery

Richard Owen showed that fossils of extinct species Darwin found in South America were allied to living species on the same continent. In March 1837, ornithologist John Gould announced that Darwin's Rhea was a separate species from the previously described rhea (though their territories overlapped), HP Pavilion N5100 Battery

that mockingbirds collected on the Galápagos Islands represented three separate species each unique to a particular island, and that several distinct birds from those islands were all classified as finches.[22] Darwin began speculating, in a series of notebooks, HP Pavilion N5200 Battery

on the possibility that "one species does change into another" to explain these findings, and around July sketched a genealogical branching of a single evolutionary tree, discarding Lamarck's independent lineages progressing to higher forms.[23] HP Pavilion N5250 Battery

Unconventionally, Darwin asked questions of fancy pigeon and animal breeders as well as established scientists. At the zoo he had his first sight of an ape, and was profoundly impressed by how human the orangutan seemed.[24]HP Pavilion N5290 Battery

In late September 1838, he started reading Thomas Malthus's An Essay on the Principle of Population with its statistical proof that human populations breed beyond their means and struggle to survive. Darwin related this to the struggle for existence among wildlife and botanist de Candolle's "warring of the species" in plants; HP Pavilion N5300 Battery

he immediately envisioned "a force like a hundred thousand wedges" pushing well-adapted variations into "gaps in the economy of nature", so that the survivors would pass on their form and abilities, and unfavourable variations would be destroyed.[25][26] HP Pavilion N5340 Battery

By December 1838, he had noted a similarity between the act of breeders selecting traits and a Malthusian Nature selecting among variants thrown up by "chance" so that "every part of newly acquired structure is fully practical and perfected".[27]HP Pavilion N5350 Battery

Darwin now had the framework of his theory of natural selection "by which to work",[28] but he was fully occupied with his career as a geologist and held off writing a sketch of his theory until his book on The Structure and Distribution of Coral Reefs was completed in May 1842.[29][30]HP Pavilion N5390 Battery

FURTHER DEVELOPMENT

Darwin continued to research and extensively revise his theory while focusing on his main work of publishing the scientific results of the Beagle voyage.[29] He tentatively wrote of his ideas to Lyell in January 1842;[31] then in June he roughed out a 35-page "Pencil Sketch" of his theory.[32]HP Pavilion N5400 Battery

 Darwin began correspondence about his theorising with the botanist Joseph Dalton Hooker in January 1844, and by July had rounded out his "sketch" into a 230-page "Essay", to be expanded with his research results and published if he died prematurely.[33]HP Pavilion N5415 Battery

In November 1844, the anonymously published popular science book Vestiges of the Natural History of Creation, written by Scottish journalist Robert Chambers, widened public interest in the concept of transmutation of species. HP Pavilion N5420 Battery

 Vestiges used evidence from the fossil record and embryology to support the claim that living things had progressed from the simple to the more complex over time. But it proposed a linear progression rather than the branching common descent theory behind Darwin's work in progress, HP Pavilion N5425 Battery

and it ignored adaptation. Darwin read it soon after publication, and scorned its amateurish geology and zoology,[34] but he carefully reviewed his own arguments after leading scientists, including Adam Sedgwick, attacked its morality and scientific errors.[35HP Pavilion N5430 Battery

] Vestiges had significant influence on public opinion, and the intense debate helped to pave the way for the acceptance of the more scientifically sophisticated Origin by moving evolutionary speculation into the mainstream. While few naturalists were willing to consider transmutation, Herbert Spencer became an active proponent of Lamarckism and progressive development in the 1850s.[36]HP Pavilion N5435 Battery

Hooker was persuaded to take away a copy of the "Essay" in January 1847, and eventually sent a page of notes giving Darwin much needed feedback. Reminded of his lack of expertise in taxonomy, Darwin began an eight year study of barnacles, HP Pavilion N5440 Battery

becoming the leading expert on their classification. Using his theory, he discovered homologies showing that slightly changed body parts served different functions to meet new conditions, and he found an intermediate stage in the evolution of distinct sexes.[37]HP Pavilion N5445 Battery

 

Darwin's barnacle studies convinced him that variation arose constantly and not just in response to changed circumstances. In 1854, he completed the last part of his Beagle-related writing and began working full-time on evolution. His thinking changed from the view that species formed in isolated populations only, as on islands, to an emphasis on speciation without isolation; that is, HP Pavilion N5455 Battery

he saw increasing specialisation within large stable populations as continuously exploiting new ecological niches. He conducted empirical research focusing on difficulties with his theory. He studied the developmental and anatomical differences between different breeds of many domestic animals, HP Pavilion N5470 Battery

became actively involved in fancy pigeon breeding, and experimented (with the help of his son Francis) on ways that plant seeds and animals might disperse across oceans to colonise distant islands. By 1856, his theory was much more sophisticated, with a mass of supporting evidence.[37][38]HP Pavilion N5490 Battery

EVENTS LEADING TO PUBLICATION

An 1855 paper on the "introduction" of species, written by Alfred Russel Wallace, claimed that patterns in the geographical distribution of living and fossil species could be explained if every new species always came into existence near an already existing, closely related species.[39] HP Pavilion N6000 Battery

Charles Lyell recognised the implications of Wallace's paper and its possible connection to Darwin's work, although Darwin did not, and in the spring of 1856 Lyell urged Darwin to publish his theory to establish priority. Darwin was torn between the desire to set out a full and convincing account and the pressure to quickly produce a short paper. HP Pavilion n6400 Battery

He decided he did not want to expose his ideas to review by an editor as would have been required to publish in an academic journal. On 14 May 1856, he began a "sketch" account, and by July had decided to produce a full technical treatise on species.[40]HP Pavilion N6490 Battery

Darwin was hard at work on his "big book" on Natural Selection, when on 18 June 1858 he received a parcel from Wallace, who stayed on the Maluku Islands (Ternate and Gilolo). It enclosed twenty pages describing an evolutionary mechanism, a response to Darwin's recent encouragement, HP Pavilion XF100 Battery

with a request to send it on to Lyell if Darwin thought it worthwhile. The mechanism was similar to Darwin's own theory.[40] Darwin wrote to Lyell that "your words have come true with a vengeance, ... forestalled" and he would "of course, at once write and offer to send [it] to any journal" that Wallace chose, adding that "all my originality, whatever it may amount to, will be smashed".[41] HP Pavilion XZ100 Battery

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Lyell and Hooker agreed that a joint paper should be presented at the Linnean Society, and on 1 July 1858, the papers entitled On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection, HP Pavilion ZD7000 Battery

by Wallace and Darwin respectively, were read out but drew little reaction. While Darwin considered Wallace's idea to be identical to his concept of natural selection, historians have pointed out differences. Darwin described natural selection as being analogous to the artificial selection practised by animal breeders, HP Pavilion ZD7900 Battery

and emphasised competition between individuals; Wallace drew no comparison to selective breeding, and focused on ecological pressures that kept different varieties adapted to local conditions.[42][43][44]HP Pavilion ZE2000 Battery

 

After the meeting, Darwin decided to write "an abstract of my whole work".[45] He started work on 20 July 1858, while on holiday atSandown,[46] and wrote parts of it from memory.[47] Lyell discussed arrangements with publisher John Murray III, of the publishing house John Murray,[48] who responded immediately to Darwin's letter of 31 March 1859[49] HP Pavilion ZT1000 Battery

with an agreement to publish the book without even seeing the manuscript, and an offer to Darwin of 23 of the profits.[50] (eventually Murray paid £180 to Darwin for the 1st edition and by Darwin's death in 1882 the book was in its 6th edition, earning Darwin nearly £3000.[51]) HP Pavilion ZT1125 Battery

Darwin had initially decided to call it An abstract of an Essay on the Origin of Species and Varieties Through natural selection, but with Murray's persuasion it was eventually changed to the snappier title: On the Origin of Species, with the title page adding by Means of Natural Selection, HP Pavilion ZT1130 Battery

or the Preservation of Favoured Races in the Struggle for Life.[1] Here the term "races" is used as an alternative for "varieties"and does not carry the modern connotation of human races—the first use in the book refers to "the several races, for instance, of the cabbage" and proceeds to a discussion of "the hereditary varieties or races of our domestic animals and plants".[52]HP Pavilion ZT1145 Battery

TIME TAKEN TO PUBLISH

Darwin had his basic theory of natural selection "by which to work" by December 1838, yet almost twenty years later, when Wallace's letter arrived on 18 June 1858, Darwin was still not ready to publish his theory. It was long thought that Darwin avoided or delayed making his ideas public for personal reasons. HP Pavilion ZT1150 Battery

Reasons suggested have included fear of religious persecution or social disgrace if his views were revealed, and concern about upsetting his clergymen naturalist friends or his pious wife Emma. Charles Darwin's illness caused repeated delays. HP Pavilion ZT1155 Battery

His paper on Glen Roy had proved embarrassingly wrong, and he may have wanted to be sure he was correct. David Quammen has suggested all these factors may have contributed, and notes Darwin's large output of books and busy family life during that time.[53]HP Pavilion ZT1160 Battery

A more recent study by science historian John van Wyhe has determined that the idea that Darwin delayed publication only dates back to the 1940s, and Darwin's contemporaries thought the time he took was reasonable. HP Pavilion ZD7300 Battery

Darwin always finished one book before starting another. While he was researching, he told many people about his interest in transmutation without causing outrage. He firmly intended to publish, but it was not until September 1854 that he could work on it full time. His estimate that writing his "big book" would take five years was optimistic.[54]HP Pavilion ZT1170 Battery

PUBLICATION AND SUBSEQUENT EDITIONS

On the Origin of Species was first published on Thursday 24 November 1859, priced at fifteen shillings. The book had been offered to booksellers at Murray's autumn sale on Tuesday 22 November, and all available copies had been taken up immediately. HP Pavilion ZT1175 Battery

In total, 1,250 copies were printed but after deducting presentation and review copies, and five for Stationers' Hall copyright, around 1,170 copies were available for sale.[1] Significantly, 500 were taken by Mudie's Library, ensuring that the book promptly reached a large number of subscribers to the library.[55] The second edition of 3,000 copies was quickly brought out on 7 January 1860,[56]HP Pavilion ZT1180 Battery

 and incorporated numerous corrections as well as a response to religious objections by the addition of a new epigraph on page ii, a quotation fromCharles Kingsley, and the phrase "by the Creator" amended to the closing sentence.[57] HP Pavilion ZT1185 Battery

During Darwin's lifetime the book went through six editions, with cumulative changes and revisions to deal with counter-arguments raised. The third edition came out in 1861, with a number of sentences rewritten or added and an introductory appendix, HP Pavilion ZT1190 Battery

An Historical Sketch of the Recent Progress of Opinion on the Origin of Species,[58] while the fourth in 1866 had further revisions. The fifth edition, published on 10 February 1869, incorporated more changes and for the first time included the phrase "survival of the fittest", which had been coined by the philosopher Herbert Spencer in his Principles of Biology (1864).[59]HP Pavilion ZT1195 Battery

 

In January 1871, George Jackson Mivart's On the Genesis of Species listed detailed arguments against natural selection, and claimed it included falsemetaphysics.[60] Darwin made extensive revisions to the sixth edition of the Origin (this was the first edition in which he used the word "evolution"[61]), and added a new chapter VII, Miscellaneous objections, to address Mivart's arguments.[62] HP Pavilion ZT1200 Battery

The sixth edition was published by Murray on 19 February 1872 with "On" dropped from the title. Darwin had told Murray of working men in Lancashire clubbing together to buy the 5th edition at fifteen shillings and wanted it made more widely available; the price was halved to 7s 6d by printing in a smaller font. It includes a glossary compiled by W.S. Dallas. Book sales increased from 60 to 250 per month.[62]HP Pavilion ZT1210 Battery

 

PUBLICATION OUTSIDE GREAT BRITAIN

In the United States, Asa Gray negotiated with a Boston publisher for publication of an authorised American version, but learnt that two New York publishing firms were already planning to exploit the absence of international copyright to printOriginHP Pavilion ZT1230 Battery

Darwin was delighted by the popularity of the book, and asked Gray to keep any profits.[64] Gray managed to negotiate a 5% royalty with Appleton's of New York,[65] who got their edition out in mid January 1860, and the other two withdrew. In a May letter, Darwin mentioned a print run of 2,500 copies, but it is not clear if this referred to the first printing only as there were four that year.[1][66]HP Pavilion ZT1250 Battery

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The book was widely translated in Darwin's lifetime, but problems arose with translating concepts and metaphors, and some translations were biased by the translator's own agenda.[67] Darwin distributed presentation copies in France and Germany, hoping that suitable applicants would come forward, HP Pavilion ZT1290 Battery

as translators were expected to make their own arrangements with a local publisher. He welcomed the distinguished elderly naturalist and geologist Heinrich Georg Bronn, but the German translation published in 1860 imposed Bronn's own ideas, adding controversial themes that Darwin had deliberately omitted. Bronn translated "favoured races" as "perfected races",HP Pavilion ZT1270 Battery

and added essays on issues including the origin of life, as well as a final chapter on religious implications partly inspired by Bronn's adherence to Naturphilosophie.[68] In 1862, Bronn produced a second edition based on the third English edition and Darwin's suggested additions, HP Pavilion ZT3000 Battery

but then died of a heart attack.[69]Darwin corresponded closely with Julius Victor Carus, who published an improved translation in 1867.[70] Darwin's attempts to find a translator in France fell through, and the translation by Clémence Royer published in 1862 added an introduction praising Darwin's ideas as an alternative to religious revelation and promoting ideas anticipating social Darwinism and eugenics, HP Pavilion ZT4000 Battery

as well as numerous explanatory notes giving her own answers to doubts that Darwin expressed. Darwin corresponded with Royer about a second edition published in 1866 and a third in 1870, but he had difficulty getting her to remove her notes and was troubled by these editions.[69][71] He remained unsatisfied until a translation by Edmond Barbier was published in 1876.[1] HP Pavilion ZV5000 Battery

A Dutch translation by Tiberius Cornelis Winkler was published in 1860.[72] By 1864, additional translations had appeared in Italian and Russian.[67] In Darwin's lifetime, Origin was published in Swedish in 1871,[73] Danish in 1872, Polish in 1873, Hungarian in 1873–1874, Spanish in 1877 and Serbian in 1878. By 1977, it had appeared in an additional 18 languages.[74]HP Pavilion ZV5200 Battery

TITLE PAGES AND INTRODUCTION

Page ii contains quotations by William Whewell and Francis Bacon on the theology of natural laws,[75] harmonising science and religion in accordance with Isaac Newton's belief in a rational God who established a law-abiding cosmos.[76] In the second edition, Darwin added an epigraph from Joseph Butler affirming that God could work through scientific laws as much as throughmiracles, HP Pavilion ZV6000 Battery

in a nod to the religious concerns of his oldest friends.[57] The Introduction establishes Darwin's credentials as a naturalist and author,[77] then refers to John Herschel's letter suggesting that the origin of species "would be found to be a natural in contradistinction to a miraculous process":[78]HP Pavilion ZX5000 Battery

 

WHEN on board H.M.S. 'Beagle,' as naturalist, I was much struck with certain facts in the distribution of the inhabitants of South America, and in the geological relations of the present to the past inhabitants of that continent. These facts seemed to me to throw some light on the origin of species—that mystery of mysteries, as it has been called by one of our greatest philosophers.[79]Sony vaio VPCCW2S1E/B Battery

 

Darwin refers specifically to the distribution of the species rheas, and to that of the Galápagos tortoises and mockingbirds. He mentions his years of work on his theory, and the arrival of Wallace at the same conclusion, which led him to "publish this Abstract" of his incomplete work. He outlines his ideas, and sets out the essence of his theory: Sony vaio VPCCW2S1E/L Battery

 

As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, Sony vaio VPCCW2S1E/P Battery

under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be naturally selected. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.[80]Sony vaio VPCCW2S1E/R Battery

 

Starting with the third edition, Darwin prefaced the introduction with a sketch of the historical development of evolutionary ideas.[81] In that sketch he acknowledged that Patrick Matthew had, unknown to Wallace or himself, anticipated the concept of natural selection in an appendix to a book published in 1831;[82] in the fourth edition he mentioned that William Charles Wells had done so as early as 1813.[83]Sony vaio VPCCW2S1E/W Battery

 

VARIATION UNDER DOMESTICATION AND UNDER NATURE

Chapter I covers animal husbandry and plant breeding, going back to ancient Egypt. Darwin discusses contemporary opinions on the origins of different breeds under cultivation to argue that many have been produced from common ancestors by selective breeding.[84] HP Pavilion ZX5200 Battery

As an illustration of artificial selection, he describesfancy pigeon breeding,[85] noting that "[t]he diversity of the breeds is something astonishing", yet all were descended from one species of rock pigeon.[86]Darwin saw two distinct kinds of variation: (1) Sony VPCB11V9E/B Battery

rare abrupt changes he called "sports" or "monstrosities" (example: ancon sheep with short legs), and (2) ubiquitous small differences (example: slightly shorter or longer bill of pigeons).[87] Both types of hereditary changes can be used by breeders. However, for Darwin the small changes were most important in evolution. Sony VPCB11X9E/B Battery

In Chapter II, Darwin specifies that the distinction between species and varieties is arbitrary, with experts disagreeing and changing their decisions when new forms were found. He concludes that "a well-marked variety may be justly called an incipient species" and that "species are only strongly marked and permanent varieties".[88] He argues for the ubiquity of variation in nature.[89]Sony VPCF11C4E/B Battery

 Historians have noted that naturalists had long been aware that the individuals of a species differed from one another, but had generally considered such variations to be limited and unimportant deviations from the archetype of each species, that archetype being a fixed ideal in the mind of God. Darwin and Wallace made variation among individuals of the same species central to understanding the natural world.[85]Sony VPCF11C5E Battery

 

STRUGGLE FOR EXISTENCE, NATURAL SELECTION, AND DIVERGENCE

In Chapter III, Darwin asks how varieties "which I have called incipient species" become distinct species, and in answer introduces the key concept he calls "natural selection";[90] in the fifth edition he adds, "But the expression often used by Mr. Herbert Spencer, of the Survival of the Fittest, is more accurate, and is sometimes equally convenient."[91]Sony VPCF11D4E Battery

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Owing to this struggle for life, any variation, however slight and from whatever cause proceeding, if it be in any degree profitable to an individual of any species, in its infinitely complex relations to other organic beings and to external nature, Sony vaio VPCF12F4E/H battery

will tend to the preservation of that individual, and will generally be inherited by its offspring ... I have called this principle, by which each slight variation, if useful, is preserved, by the term of Natural Selection, in order to mark its relation to man's power of selection.[90]Sony vaio VPCF12M1E/H battery

 

He notes that both A. P. de Candolle and Charles Lyell had stated that all organisms are exposed to severe competition. Darwin emphasizes that he used the phrase "struggle for existence" in "a large and metaphorical sense, including dependence of one being on another";Sony vaio VPCF12S1E/B battery

he gives examples ranging from plants struggling against drought to plants competing for birds to eat their fruit and disseminate their seeds. He describes the struggle resulting from population growth: "It is the doctrine of Malthus applied with manifold force to the whole animal and vegetable kingdoms." Sony vaio VPCF12Z1E/BI battery

He discusses checks to such increase including complex ecological interdependencies, and notes that competition is most severe between closely related forms "which fill nearly the same place in the economy of nature".[92]Sony vaio VPCF13E4E battery

 

Chapter IV details natural selection under the "infinitely complex and close-fitting ... mutual relations of all organic beings to each other and to their physical conditions of life".[93] Darwin takes as an example a country where a change in conditions led to extinction of some species, Sony vaio VPCF13E8E/H battery

immigration of others and, where suitable variations occurred, descendants of some species became adapted to new conditions. He remarks that the artificial selection practised by animal breeders frequently produced sharp divergence in character between breeds, and suggests that natural selection might do the same, saying: Sony vaio VPCF13J0E/H battery

But how, it may be asked, can any analogous principle apply in nature? I believe it can and does apply most efficiently, from the simple circumstance that the more diversified the descendants from any one species become in structure, constitution, and habits, by so much will they be better enabled to seize on many and widely diversified places in the polity of nature, and so be enabled to increase in numbers.[94]Sony vaio VPCF13L8E/H battery

Historians have remarked that here Darwin anticipated the modern concept of an ecological niche.[95] He did not suggest that every favourable variation must be selected, nor that the favoured animals were better or higher, but merely more adapted to their surroundings. Sony vaio VPCF13M1E/B battery Darwin proposes sexual selection, driven by competition between males for mates, to explain sexually dimorphic features such as lion manes, deer antlers, peacock tails, bird songs, and the bright plumage of some male birds.[96] Sony vaio VPCF13M1E/B battery

He analysed sexual selection more fully in The Descent of Man, and Selection in Relation to Sex (1871). Natural selection was expected to work very slowly in forming new species, but given the effectiveness of artificial selection, he could "see no limit to the amount of change, to the beauty and infinite complexity of the coadaptations between all organic beings, Sony vaio VPCF13M1E/H battery

one with another and with their physical conditions of life, which may be effected in the long course of time by nature's power of selection". Using a tree diagram and calculations, he indicates the "divergence of character" from original species into new species and genera. He describes branches falling off as extinction occurred, while new branches formed in "the great Tree of life ... with its ever branching and beautiful ramifications".[97]Sony vaio VPCF13S1E/B battery

 

VARIATION AND HEREDITY

In Darwin's time there was no agreed-upon model of heredity;[98] in Chapter I Darwin admitted, "The laws governing inheritance are quite unknown."[99] He accepted a version of the inheritance of acquired characteristics (which after Darwin's death came to be called Lamarckism), Sony vaio VPCF13S8E battery

and Chapter V discusses what he called the effects of use and disuse; he wrote that he thought "there can be little doubt that use in our domestic animals strengthens and enlarges certain parts, and disuse diminishes them; and that such modifications are inherited", Sony vaio VPCF13S8E/B battery

and that this also applied in nature.[100] Darwin stated that some changes that were commonly attributed to use and disuse, such as the loss of functional wings in some island dwelling insects, might be produced by natural selection. In later additions of Origin, Sony vaio VPCF13V5002B battery

Darwin expanded the role attributed to the inheritance of acquired characteristics. Darwin also admitted ignorance of the source of inheritable variations, but speculated they might be produced by environmental factors.[101][102] However, one thing was clear: Sony vaio VPCF13Z1E/B battery

whatever the exact nature and causes of new variations, Darwin knew from observation and experiment that breeders were able to select such variations and produce huge differences in many generations of selection.[87] The observation that selection works in domestic animals is not destroyed by lack of understanding of the underlying hereditary mechanism. Sony vaio VPCF13Z8E/BI battery

 

Breeding of animals and plants showed related varieties varying in similar ways, or tending to revert to an ancestral form, and similar patterns of variation in distinct species were explained by Darwin as demonstrating common descent. Sony vaio VPCF21Z1E battery

He recounted how Lord Morton's mare apparently demonstrated telegony, offspring inheriting characteristics of a previous mate of the female parent, and accepted this process as increasing the variation available for natural selection.[103]Sony vaio VPCF21Z1E/BI battery

 

More detail was given in Darwin's 1868 book on The Variation of Animals and Plants under Domestication, which tried to explain heredity through his hypothesis of pangenesis. Although Darwin had privately questioned blending inheritance,Sony vaio VPCF22J1E battery

he struggled with the theoretical difficulty that novel individual variations would tend to blend into a population. However, inherited variation could be seen,[104] and Darwin's concept of selection working on a population with a range of small variations was workable.[105] It was not until the modern evolutionary synthesis in the 1930s and 1940s that a model of heredity became completely integrated with a model of variation.[106]Sony vaio VPCF22L1E battery

DIFFICULTIES FOR THE THEORY

Chapter VI begins by saying the next three chapters will address possible objections to the theory, the first being that often no intermediate forms between closely related species are found, though the theory implies such forms must have existed.Sony vaio VPCF22M1E battery

Darwin attributed this to the competition between different forms, combined with the small number of individuals of intermediate forms, often leading to extinction of such forms.[107] The rest of the chapter deals with whether natural selection could produce complex specialised structures, and the behaviours to use them, when it would be difficult to imagine how intermediate forms could be functional. Darwin said: Sony vaio VPCF22S1E battery

 

Secondly, is it possible that an animal having, for instance, the structure and habits of a bat, could have been formed by the modification of some animal with wholly different habits? Can we believe that natural selection could produce, Sony vaio VPCF22S8E battery

on the one hand, organs of trifling importance, such as the tail of a giraffe, which serves as a fly-flapper, and, on the other hand, organs of such wonderful structure, as the eye, of which we hardly as yet fully understand the inimitable perfection?[108]Sony vaio VPCF23A9E battery

 

His answer was that in many cases animals exist with intermediate structures that are functional. He presented flying squirrels, and flying lemurs as examples of how bats might have evolved from non-flying ancestors.[109] Sony vaio VPCF23M1E battery

He discussed various simple eyes found in invertebrates, starting with nothing more than an optic nerve coated with pigment, as examples of how the vertebrate eye could have evolved. Darwin concludes: "If it could be demonstrated that any complex organ existed, which could not possibly have been formed by numerous, successive, slight modifications, my theory would absolutely break down. But I can find out no such case."[110]Sony vaio VPCF23Q1E battery

 

Chapter VII (of the first edition) addresses the evolution of instincts. His examples included two he had investigated experimentally: slave-making ants and the construction of hexagonal cells by honey bees. Darwin noted that some species of slave-making ants were more dependent on slaves than others, Sony vaio VPCF23S1E battery

and he observed that many ant species will collect and store the pupae of other species as food. He thought it reasonable that species with an extreme dependency on slave workers had evolved in incremental steps. He suggested that bees that make hexagonal cells evolved in steps from bees that made round cells, under pressure from natural selection to economise wax. Darwin concluded: Sony vaio VPCF23S1E battery

Finally, it may not be a logical deduction, but to my imagination it is far more satisfactory to look at such instincts as the young cuckoo ejecting its foster-brothers,Sony vaio VPCF24Q1E battery

 —ants making slaves, —the larvæ of ichneumonidæ feeding within the live bodies of caterpillars, —not as specially endowed or created instincts, but as small consequences of one general law, leading to the advancement of all organic beings, namely, multiply, vary, let the strongest live and the weakest die.[111]Sony vaio VPCF24M1E battery

 

Chapter VIII addresses the idea that species had special characteristics that prevented hybrids from being fertile in order to preserve separately created species. Darwin said that, far from being constant, the difficulty in producing hybrids of related species, Sony VPCCA2S1E Battery

and the viability and fertility of the hybrids, varied greatly, especially among plants. Sometimes what were widely considered to be separate species produced fertile hybrid offspring freely, and in other cases what were considered to be mere varieties of the same species could only be crossed with difficulty.Sony VPCCA2S1E/D.G4 Battery

Darwin concluded: "Finally, then, the facts briefly given in this chapter do not seem to me opposed to, but even rather to support the view, that there is no fundamental distinction between species and varieties."[112]Sony VPCCA2S1E/L Battery

In the sixth edition Darwin inserted a new chapter VII (renumbering the subsequent chapters) to respond to criticisms of earlier editions, including the objection that many features of organisms were not adaptive and could not have been produced by natural selection. Sony VPCCA2S1E/R Battery

He said some such features could have been by-products of adaptive changes to other features, and that often features seemed non-adaptive because their function was unknown, as shown by his book onFertilisation of Orchids that explained how their elaborate structures facilitated pollination by insects. Sony VPCCA3S1E Battery

Much of the chapter responds to George Jackson Mivart's criticisms, including his claim that features such as baleen filters in whales, flatfish with both eyes on one side and the camouflage of stick insectscould not have evolved through natural selection because intermediate stages would not have been adaptive. Darwin proposed scenarios for the incremental evolution of each feature.[113]Sony VPCCB2S1E Battery

 

GEOLOGIC RECORD

Chapter IX deals with the fact that the geologic record appears to show forms of life suddenly arising, without the innumerable transitional fossils expected from gradual changes. Darwin borrowed Charles Lyell's argument in Principles of Geology that the record is extremely imperfect as fossilisation is a very rare occurrence, Sony VPCCB2S1E/B Battery

spread over vast periods of time; since few areas had been geologically explored, there could only be fragmentary knowledge of geological formations, and fossil collections were very poor. Evolved local varieties which migrated into a wider area would seem to be the sudden appearance of a new species. Sony VPCCB2S1E/W Battery

Darwin did not expect to be able to reconstruct evolutionary history, but continuing discoveries gave him well founded hope that new finds would occasionally reveal transitional forms.[114][115] To show that there had been enough time for natural selection to work slowly, Sony VPCCB3S1E Battery

he again cited Principles of Geology and other observations based on sedimentation and erosion, including an estimate that erosion of The Weald had taken 300 million years.[116] The initial appearance of entire groups of well developed organisms in the oldest fossil-bearing layers, Sony VPCCB3S1E/B Battery

now known as the Cambrian explosion, posed a problem. Darwin had no doubt that earlier seas had swarmed with living creatures, but stated that he had no satisfactory explanation for the lack of fossils.[117] Fossil evidence ofpre-Cambrian life has since been found, extending the history of life back for billions of years.[118]Sony VPCCB3S1E/W Battery

 

Chapter X examines whether patterns in the fossil record are better explained by common descent and branching evolution through natural selection than by the individual creation of fixed species. Darwin expected species to change slowly, but not at the same rate – some organisms such as Lingula were unchanged since the earliest fossils Sony VPCCB4Z1E Battery

. The pace of natural selection would depend on variability and change in the environment.[119] This distanced his theory from Lamarckianlaws of inevitable progress.[114] It has been argued that this anticipated the punctuated equilibrium hypothesis,[115][120]Sony VPCCB4Z1E/B Battery

 but other scholars have preferred to emphasise Darwin's commitment to gradualism.[121] He cited Richard Owen's findings that the earliest members of a class were a few simple and generalised species with characteristics intermediate between modern forms, and were followed by increasingly diverse and specialised forms, matching the branching of common descent from an ancestor.[114] Sony VPCCB4Z1E/W Battery

Patterns of extinction matched his theory, with related groups of species having a continued existence until extinction, then not reappearing. Recently extinct species were more similar to living species than those from earlier eras, and as he had seen in South America, and William Clifthad shown in Australia, fossils from recent geological periods resembled species still living in the same area.[119]Sony VPCCB4Z1E/W.G4 Battery

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GEOGRAPHIC DISTRIBUTION

Chapter XI deals with evidence from biogeography, starting with the observation that differences in flora and fauna from separate regions cannot be explained by environmental differences alone; South America, Africa, and Australia all have regions with similar climates at similar latitudes, //www.orderbattery.co.uk/sony-vaio-vpcy11s1e-s-battery.html">Sony vaio VPCY11S1E/S battery

but those regions have very different plants and animals. The species found in one area of a continent are more closely allied with species found in other regions of that same continent than to species found on other continents. Sony vaio VPCY21S1E/G battery

Darwin noted that barriers to migration played an important role in the differences between the species of different regions. The coastal sea life of the Atlantic and Pacific sides of Central America had almost no species in common even though the Isthmus of Panama was only a few miles wide. His explanation was a combination of migration and descent with modification. Sony vaio VPCY21S1E/L battery

He went on to say: "On this principle of inheritance with modification, we can understand how it is that sections of genera, whole genera, and even families are confined to the same areas, as is so commonly and notoriously the case."[122] Sony vaio VPCY21S1E/SI battery

Darwin explained how a volcanic island formed a few hundred miles from a continent might be colonised by a few species from that continent. These species would become modified over time, Sony vaio VPCY22Z5008B battery

but would still be related to species found on the continent, and Darwin observed that this was a common pattern. Darwin discussed ways that species could be dispersed across oceans to colonise islands, many of which he had investigated experimentally.[123]Sony vaio VPCYA1V9E/B battery

 

Chapter XII continues the discussion of biogeography. After a brief discussion of freshwater species, it returns to oceanic islands and their peculiarities; for example on some islands roles played by mammals on continents were played by other animals such as flightless birds or reptiles. The summary of both chapters says: Sony vaio VPCYB1S1E/S battery

 

... I think all the grand leading facts of geographical distribution are explicable on the theory of migration (generally of the more dominant forms of life), together with subsequent modification and the multiplication of new forms. Sony PCG-81111T battery 

We can thus understand the high importance of barriers, whether of land or water, which separate our several zoological and botanical provinces. We can thus understand the localisation of sub-genera, genera, and families; and how it is that under different latitudes, Sony vaio VPCYA15FH/R battery

for instance in South America, the inhabitants of the plains and mountains, of the forests, marshes, and deserts, are in so mysterious a manner linked together by affinity, and are likewise linked to the extinct beings which formerly inhabited the same continent ...  Sony vaio VPCS13L8E/B battery

On these same principles, we can understand, as I have endeavoured to show, why oceanic islands should have few inhabitants, but of these a great number should be endemic or peculiar; ...[124] Sony PCG-5J1L battery

 

CLASSIFICATION, MORPHOLOGY, EMBRYOLOGY, RUDIMENTARY ORGANS

Chapter XIII starts by observing that classification depends on species being grouped together in a multilevel system of groups and sub groups based on varying degrees of resemblance. After discussing classification issues, Darwin concludes:  Dell Inspiron 15z battery

 

All the foregoing rules and aids and difficulties in classification are explained, if I do not greatly deceive myself, on the view that the natural system is founded on descent with modification; that the characters which naturalists consider as showing true affinity between any two or more species, Sony PCG-5L1L battery

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are those which have been inherited from a common parent, and, in so far, all true classification is genealogical; that community of descent is the hidden bond which naturalists have been unconsciously seeking, ...[125]Sony VGN-CR42Z/R battery

 

Darwin discusses morphology, including the importance of homologous structures. He says, "What can be more curious than that the hand of a man, formed for grasping, that of a mole for digging, the leg of the horse, the paddle of the porpoise, and the wing of the bat, should all be constructed on the same pattern, and should include the same bones, in the same relative positions?"[126] Sony VPCB11V9E/B Battery

He notes that animals of the same class often have extremely similar embryos. Darwin discusses rudimentary organs, such as the wings of flightless birds and the rudiments of pelvis and leg bones found in some snakes. He remarks that some rudimentary organs, such as teeth in baleen whales, are found only in embryonic stages. Sony VPCB11X9E/B Battery

 

CONCLUDING REMARKS

The final chapter reviews points from earlier chapters, and Darwin concludes by hoping that his theory might produce revolutionary changes in many fields of natural history. Although he avoids the controversial topic of human origins in the rest of the book so as not to prejudice readers against his theory, Sony VPCF11C4E/B Battery

here he ventures a cautious hint that psychology would be put on a new foundation and that "Light will be thrown on the origin of man".[127] Darwin ends with a passage that became well known and much quoted: Sony VPCF11C5E Battery

 

It is interesting to contemplate an entangled bank, clothed with many plants of many kinds, with birds singing on the bushes, with various insects flitting about, and with worms crawling through the damp earth, and to reflect that these elaborately constructed forms, so different from each other, and dependent on each other in so complex a manner, Sony VPCF11D4E Battery

have all been produced by laws acting around us ... Thus, from the war of nature, from famine and death, the most exalted object which we are capable of conceiving, namely, the production of the higher animals, directly follows. There is grandeur in this view of life, Sony vaio VPCF11M1E/H battery

with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.[128]Sony vaio VPCF12F4E/H battery

 

NATURE AND STRUCTURE OF DARWIN'S ARGUMENT

Darwin's aims were twofold: to show that species had not been separately created, and to show that natural selection had been the chief agent of change.[129] He knew that his readers were already familiar with the concept of transmutation of species from Vestiges, and his introduction ridicules that work as failing to provide a viable mechanism.[130] Sony vaio VPCF12M1E/H battery

Therefore the first four chapters lay out his case that selection in nature, caused by the struggle for existence, is analogous to the selection of variations under domestication, and that the accumulation of adaptive variations provides a scientifically testable mechanism for evolutionary speciation.[131][132]Sony vaio VPCF12S1E battery

Later chapters provide evidence that evolution has occurred, supporting the idea of branching, adaptive evolution without directly proving that selection is the mechanism. Darwin presents supporting facts drawn from many disciplines, Sony vaio VPCF12S1E/B battery

showing that his theory could explain a myriad of observations from many fields of natural history that were inexplicable under the alternate concept that species had been individually created.[132][133][134] The structure of Darwin's argument showed the influence of John Herschel, Sony vaio VPCF12Z1E/BI battery

whose philosophy of science maintained that a mechanism could be called a vera causa (true cause) if three things could be demonstrated: its existence in nature, its ability to produce the effects of interest, and its ability to explain a wide range of observations.[135]Sony vaio VPCF13E4E battery

 

LITERARY STYLE

The Examiner review of 3 December 1859 commented, "Much of Mr. Darwin's volume is what ordinary readers would call 'tough reading;' that is, writing which to comprehend requires concentrated attention and some preparation for the task. All, however, is by no means of this description, and many parts of the book abound in information, easy to comprehend and both instructive and entertaining."[130][136]Sony vaio VPCF13E8E/H battery

 

While the book was readable enough to sell, its dryness ensured that it was seen as aimed at specialist scientists and could not be dismissed as mere journalism or imaginative fiction. Unlike the still-popular Vestiges, it avoided the narrative style of the historical novel and cosmological speculation, though the closing sentence clearly hinted at cosmic progression. Sony vaio VPCF13J0E/H battery

Darwin had long been immersed in the literary forms and practices of specialist science, and made effective use of his skills in structuring arguments.[130] David Quammen has described the book as written in everyday language for a wide audience, but noted that Darwin's literary style was uneven: in some places he used convoluted sentences that are difficult to read; Sony vaio VPCF13L8E/H battery

in other places his writing was beautiful. Quammen advised that later editions were weakened by Darwin making concessions and adding details to address his critics, and recommended the first edition.[137]James T. Costa said that because the book was an abstract produced in haste in response to Wallace's essay, Sony vaio VPCF13M1E/B battery

it was more approachable than the big book on natural selection Darwin had been working on, which would have been encumbered by scholarly footnotes and much more technical detail. He added that parts ofOrigin are dense, but parts are almost lyrical, and the case studies and observations are presented in a narrative style unusual in serious scientific books, which broadened its audience.[138]Sony vaio VPCF13M1E/H battery

 

Reception

The book aroused international interest[140] and a widespread debate, with no sharp line between scientific issues and ideological, social and religious implications.[141] Much of the initial reaction was hostile, but Darwin had to be taken seriously as a prominent and respected name in science. There was much less controversy than had greeted the 1844 publicationVestiges of Creation, Sony vaio VPCF13S1E/B battery

which had been rejected by scientists,[140] but had influenced a wide public readership into believing that nature and human society were governed by natural laws.[25] The Origin of Species as a book of wide general interest became associated with ideas of social reform. Sony vaio VPCF13S8E battery

Its proponents made full use of a surge in the publication of review journals, and it was given more popular attention than almost any other scientific work, though it failed to match the continuing sales of Vestiges.[142] Sony vaio VPCF13S8E/B battery

Darwin's book legitimised scientific discussion of evolutionary mechanisms, and the newly coined termDarwinism was used to cover the whole range of evolutionism, not just his own ideas. By the mid 1870s, evolutionism was triumphant.[141]Sony vaio VPCF13V5002B battery

With the exception of a brief hint in the final chapter, Darwin had avoided the subject of human evolution. Despite this, the first review claimed it made a creed of the "men from monkeys" idea from Vestiges.[143] Human evolution became central to the debate and was strongly argued by Huxley who featured it in his popular "working-men's lectures". Sony vaio VPCF13Z1E/B battery

Darwin did not publish his own views on this until 1871.[144] The naturalism of natural selection conflicted with presumptions of purpose in nature and while this could be reconciled by theistic evolution, other mechanisms implying more progress or purpose were more acceptable. Sony VPCCA2S1E Battery

 Herbert Spencer had already incorporated Lamarckism into his popular philosophy of progressive free market human society. He popularised the terms evolution and survival of the fittest, and many thought Spencer was central to evolutionary thinking.[145]Sony VPCCA2S1E/D.G4 Battery

IMPACT ON THE SCIENTIFIC COMMUNITY

Scientific readers were already aware of arguments that species changed through processes that were subject to laws of nature, but the transmutational ideas of Lamarck and the vague "law of development" of Vestiges had not found scientific favour. Sony VPCCA2S1E/L Battery

Darwin presented natural selection as a scientifically testable mechanism while accepting that other mechanisms such as inheritance of acquired characters were possible. His strategy established that evolution through natural laws was worthy of scientific study, Sony VPCCA2S1E/R Battery

and by 1875, most scientists accepted that evolution occurred but few thought natural selection was significant. Darwin's scientific method was also disputed, with his proponents favouring the empiricism of John Stuart Mill's A System of Logic, Sony VPCCA3S1E Battery

while opponents held to the idealist school of William Whewell's Philosophy of the Inductive Sciences, in which investigation could begin with the intuitive truth that species were fixed objects created by design.[146] Early support for Darwin's ideas came from the findings of field naturalists studying biogeography and ecology, Sony VPCCB2S1E Battery

including Joseph Dalton Hooker in 1860, and Asa Gray in 1862.Henry Walter Bates presented research in 1861 that explained insect mimicry using natural selection. Alfred Russel Wallace discussed evidence from his Malay archipelago research, including an 1864 paper with an evolutionary explanation for the Wallace line.[147]Sony VPCCB2S1E/B Battery

 

Evolution had less obvious applications to anatomy and morphology, and at first had little impact on the research of the anatomist Thomas Henry Huxley.[149] Despite this, Huxley strongly supported Darwin on evolution; Sony VPCCB2S1E/W Battery

though he called for experiments to show whether natural selection could form new species, and questioned if Darwin'sgradualism was sufficient without sudden leaps to cause speciation. Huxley wanted science to be secular, without religious interference, Sony VPCCB3S1E Battery

and his article in the April 1860 Westminster Review promoted scientific naturalism over natural theology,[150][151]

 praising Darwin for "extending the domination of Science over regions of thought into which she has, as yet, hardly penetrated" and coining the term "Darwinism" as part of his efforts to secularise and professionalise science.[152] Huxley gained influence, and initiated the X Club, Sony VPCCB3S1E/B Battery

which used the journal Natureto promote evolution and naturalism, shaping much of late Victorian science. Later, the German morphologist Ernst Haeckel would convince Huxley that comparative anatomy and palaeontology could be used to reconstruct evolutionary genealogies.[149][153]Sony VPCCB3S1E/W Battery

 

The leading naturalist in Britain was the anatomist Richard Owen, an idealist who had shifted to the view in the 1850s that the history of life was the gradual unfolding of a divine plan.[154] Owen's review of the Origin in the April 1860 Edinburgh Review bitterly attacked Huxley, Hooker and Darwin, but also signalled acceptance of a kind of evolution as a teleological plan in a continuous "ordained becoming", Sony VPCCB4Z1E Battery

with new species appearing by natural birth. Others that rejected natural selection, but supported "creation by birth", included the Duke of Argyll who explained beauty in plumage by design.[155] Since 1858, Huxley had emphasised anatomical similarities between apes and humans, contesting Owen's view that humans were a separate sub-class. Sony VPCCB4Z1E/B Battery

Their disagreement over human origins came to the fore at the British Association for the Advancement of Science meeting featuring the legendary 1860 Oxford evolution debate.[156] In two years of acrimonious public dispute that Charles Kingsley satirised as the "Great Hippocampus Question" and parodied in The Water-Babies as the "great hippopotamus test",Sony VPCCB4Z1E/W Battery

Huxley showed that Owen was incorrect in asserting that ape brains lacked a structure present in human brains.[157] Others, including Charles Lyell and Alfred Russel Wallace, thought that humans shared a common ancestor with apes, but higher mental faculties could not have evolved through a purely material process. Darwin published his own explanation in the Descent of Man (1871).[158]Sony VPCCB4Z1E/W.G4 Battery

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